I believe it was meant as a joke to those who were expecting the eyes on the vehicle in Cars 1 to be at the headlights position. In earlier Disney films, for example Herbie, there was an anthropomorphic 1963 Volkswagen Beetle.
Though in case of Herbie, a real live action movie there was no need to animate the eyes where as in Pixar Cars much emphasis was placed.
If one were to relocate the "eyes" for Herbie,
You can see that we will lost the view of the actor inside of which there was none for Cars.
In the end, to me, it felt as an ill-intentioned attempt to be funny (but not) about why Pixar chose their location for the character's eyes.
It can even boil down to any show which places a scene which is extremely different from normally expected just for the sake of doing it
e.g. A small Simpsons scene in Family guy
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